Publications
Articles
PLOS ONE 2021 (Protocol)
Leverkus, A.B., Levy, L., Andivia, E. et al. (2021) Restoring oak forests through direct seeding or planting: Protocol for a continental-scale experiment. PLOS ONE 16(11): e0259552. https://doi.org/10.1371/journal.pone.0259552
Annals of Forest Science 2024
Medina, M., Reyes-Martín, M.P., Levy, L. et al. (2024) Ex situ germination of European acorns: data from 93 batches of 12 Quercus species. Annals of Forest Science 81, 50. https://doi.org/10.1186/s13595-024-01267-2
Datasets
Digibug Repository 2024 (Open-access)
Sampere-Medina, M., Reyes-Martín, M.P., Levy, L., Lázaro-González, A. & Leverkus, A.B. (2024) Germination of European acorns: Data from 93 batches of 12 Quercus species. [Dataset] Digibug repository of the University of Granada. https://doi.org/10.30827/Digibug.87318
Oral communications
Joint Seminar University of Freiburg 2025
Leverkus, A.B. (2025) Climate-adapted revegetation: Addressing the seeding vs. planting dilemma through a coordinated, distributed experiment. [Seminar] Chair of Silviculture and Future Forests Excellence Initiative, University of Freiburg, Germany.
Seminar Swiss Federal Research Institute WSL 2024
Leverkus, A.B. (2024) Addressing a major dilemma in assisted regeneration through a coordinated, distributed experiment: Seeding vs. planting with oaks. [Seminar] Swiss Federal Research Institute WSL, Birmensdorf, Switzerland.
Seminar Potosino Institute 2024
Leverkus, A.B. (2024) Regeneración asistida de encinos y robles: Estudios a distintas escalas para evaluar aspectos metodológicos. [Seminar] Instituto Potosino de Investigación Científica y Tecnológica, San Luis Potosí, Mexico.
Talk PEN-CAFoRR Conference 2024
Leverkus, A.B. (2024) SeedvsPlant: A Coordinated, Distributed, Continental-scale Experiment initiated at PEN-CAFoRR. [Oral communication] PEN-CAFoRR Final Conference (PEN-CAFoRR Cost Action), Belgrade, Serbia.
https://www.pen-caforr.org/wp-content/uploads/2024/10/BookofAbstracts_PEN-CAFoRRFinalConference.pdf
Talk XVI National Congress of AEET 2023
Leverkus, A.B. (2023) Regeneración por siembra o plantación: avances de un experimento a escala continental. [Oral communication] XVI Congreso Nacional de la AEET, Almería, Spain. https://congreso2023.aeet.org/resources/files/LIBRO_RESUMENES_XVI_Congreso_Nacional_de_la_AEET_2023.pdf
Talk XVI National Congress of AEET 2023
Lázaro-González, A. & Leverkus, A.B. (2023) Mapa sistemático de estudios sobre siembra y plantación para la restauración de ecosistemas. XVI Congreso Nacional de la AEET, Almería, Spain. https://congreso2023.aeet.org/resources/files/LIBRO_RESUMENES_XVI_Congreso_Nacional_de_la_AEET_2023.pdf
Scientific symposium Department of Ecology UGR 2022
Leverkus, A.B. (2022) Siembra vs. plantación de Quercus en Europa: Un experimento internacional colaborativo. [Oral communication] I Jornadas Científicas Ecología, Departament of Ecology, University of Granada, Spain.
https://ecologia.ugr.es/informacion/noticias/i-jornadas-cientificas-ecologia
Talk VIII Mexican Congress of Ecology 2022
Alexandro B. Leverkus & Laura Levy (2022) Siembra vs. plantación de Quercus en Europa: Un experimento internacional colaborativo. VIII Congreso Mexicano de Ecología, Oaxaca. https://scme.mx/wp-content/uploads/2024/02/Memorias_Congreso_Ecologia_Oax2022.pdf
Posters
Poster II CIMAS 2024
Medina M., Reyes-Martín, M.P., Merino M., Moreno-Llorca R. and Leverkus, A.B. (2024). REGENERACIÓN ASISTIDA CON QUERCÍNEAS ¿SEMBRAR O PLANTAR? UN EXPERIMENTO COLABORATIVO EUROPEO. [Poster]. II Congreso Internacional de las Montañas Sierra Nevada (CIMAS), Granada, Spain.
You can see the poster here.
Poster II CIMAS 2024
Reyes-Martín, M.P., Medina, M., Lázaro-González, A., Levy, L. and Leverkus, A.B. (2024). Collaborative assessment of seed germination for 12 european oak species. [Poster]. II Congreso Internacional de las Montañas Sierra Nevada (CIMAS), Granada, Spain.
You can see the poster here.
Poster International Oak Symposium 2024
Leverkus, A. B. (2024). Oak-Assisted Regeneration Through Seeding or Planting: A Collaborative, Continental-scale Experiment [Poster]. The International Oak Symposium: Science-based Management for Dynamic Oak Forests. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southern Research Station. https://research.fs.usda.gov/treesearch/68613
You can see the poster here.
Poster International Oak Symposium 2024
Reyes-Martín, M. P.; Sampere-Medina, M.; Lázaro-González, A.; Levy, L.; Castro, J.; Leverkus, A. B. (2024). Seed Germination of 12 European Quercus Species [Poster]. The International Oak Symposium: Science-based Management for Dynamic Oak Forests. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southern Research Station. https://research.fs.usda.gov/treesearch/68620
You can see the poster here.
Poster XVI National Congress of AEET 2023
Reyes-Martín, M.P.; Lázaro-González, A.; Moreno, M.; Levy, L. & Leverkus, A.B. (2023) Germination of Quercus seeds: test on 95 acorn batches of 12 European oak species. [Poster] XVI Congreso Nacional de la AEET, Almería, Spain. https://congreso2023.aeet.org/resources/files/LIBRO_RESUMENES_XVI_Congreso_Nacional_de_la_AEET_2023.pdf
You can see the poster here.
Poster SERE Conference 2022
Levy, L.; Solera, P.; Reyes-Martín, M.P.; Moreno, R.; Lázaro-González, A.; Rus, C.; Morales-Martín, M.; Castro, J.; Navarro, F.B. & Leverkus, A.B. (2022) Seeding vs. planting with Quercus: A collaborative, continental experiment. [Poster] Restoring Nature, Reconnecting People: SER Europe 2022 Abstract Book. https://ser-europe.org/files/2023/06/abstracts-book-2022_compressed.pdf
You can see the poster here.
Project brochures (communication with participants)
The project
Project description for participant recruitment. https://drive.google.com/file/d/1GFenRlOYG_NWZFbhIIYyPJffdVTgDtsk/view?usp=sharing
Experimental Sites
Requirements of the sites for establishing the experimental plots. Description of the experimental design and details of the experimental plots. https://drive.google.com/file/d/1aECK-zTvMxBzVtyuyodxsMKtgw2VVxPy/view?usp=sharing
Timeline & participation
Experiment timeline, tasks of participants and coordinators, required materials during the experiment, and deliverables. https://drive.google.com/file/d/1glPWmDAK0eQ6Sg2UTGd96RcQysOUSCOj/view?usp=sharing
Seeds
Specifications for acorn collection, storage, and shipment to the coordinators. Autumn 2021.
https://drive.google.com/file/d/1DeVd_8bsBqeJko3v3UdW82UkLQrYodbV/view?usp=sharing
Seeding
Assembly of the seed-shelter (seed protectors) and seeding. Reminder of the conditions of acorn collection, storage, and shipment to the coordinators. Autumn 2021 / Winter 2021-2022. https://drive.google.com/file/d/1h6wJyl7tGAcy9CBhkhDaOGhR8yBvWHmF/view?usp=sharing
Nursery, seedling selection & planting
Autumn 2022: survival assessment, selection of seedlings from the seeding treatment in the field and from the planting treatment in the nursery, growth measurements, measurements of the extracted plants (not selected), and outplanting process of the nursery seedlings to the field. https://drive.google.com/file/d/1aDiEDjzR6F517DZ9N66EoHEKhEiTXLLD/view?usp=sharing
Loggers, survival & new data system
Details on the installation of dataloggers (ambient temperature and humidity sensors) at the experimental sites. Survival assessment in spring 2023 for cohort A (one-year-old seedlings). Presentation of the new online data collection system and explanation of its functioning. Reminder about weed control. https://drive.google.com/file/d/1ntC5Rgvn78KTN5l80G3tu3FklZ94vsTX/view?usp=sharing
Survival & growth measurements - Cohort A
Growth measurements in autumn 2023 for cohort A (second growing season). Procedure for entering the data into the online data system.
https://drive.google.com/file/d/1yafuJ5Yv730qypcSCLfNfrmi5d0qNh3B/view?usp=sharing
Seedling emergence, seedling selection & planting - Cohort B
Description of the first visits for cohort B in Autumn 2023, including field emergence of seedlings from the seeding treatment, selection of seedlings from the seeding treatment in the field and from the planting treatment in the nursery, growth measurements for both treatments, description of the outplanting process for nursery seedlings in the field, and data entry into the online system. https://drive.google.com/file/d/1-2D-PJa1ODHARHp2__aFOCWVwG6rYX91/view?usp=sharing
Physiological measurements 2024 - Cohort A
Optional physiological measurements in summer 2024 for cohort A. Specifications for leaf-level gas exchange measurements with LiCor, including COâ‚‚ assimilation, Hâ‚‚O transpiration, stomatal conductance (gsw), and intercellular COâ‚‚ concentration (Ci); and for leaf water potential measurements using a Scholander chamber. https://drive.google.com/file/d/1zBsS9oT_sFTkS3k9QGzjUqteWmsw64uP/view?usp=sharing
Survival, growth measurements & seedling harvest - Cohort A
Description of the final visit for cohort A (third growing season) of the seeding vs. planting experiment in Autumn 2024, including survival assessment, growth measurements, the method for measuring Specific Leaf Area (SLA), and the harvest of seedlings for subsequent fresh and dry biomass measurements in the laboratory. Additionally, the entry of all these data into the online database.
https://drive.google.com/file/d/1dODuNjMjG7r8OiF5gL4hgW1qRuT1OqXE/view?usp=sharing
Physiological measurements 2025
Optional physiological measurements in summer 2025 for cohort B and resprouted seedlings of cohort A. Specifications for leaf-level gas exchange measurements with LiCor, including COâ‚‚ assimilation, Hâ‚‚O transpiration, stomatal conductance (gsw), and intercellular COâ‚‚ concentration (Ci); and for leaf water potential measurements using a Scholander chamber.
https://drive.google.com/file/d/111YMq4i4FkWlEZsm2DyZ_gFN1Riz89Ho/view?usp=sharing
Survival, growth measurements & seedling harvest - Cohort B
Description of the final visit for cohort B (third growing season) of the seeding vs. planting experiment in Autumn 2025, including survival assessment, growth measurements, the method for measuring Specific Leaf Area (SLA), and the harvest of seedlings for subsequent fresh and dry biomass measurements in the laboratory. Additionally, the entry of all these data into the online database.
https://drive.google.com/file/d/1147rrcl8r-YJMrDM9LnNiYcVCjjX8hvi/view?usp=sharing
Resprouting measurements - Cohort A
One year after the harvest marking the end of the seeding vs. planting experiment, an optional visit in Autumn 2025 was proposed for participants who wished to assess plant resprouting and who had at least 10 plants from each treatment (seeding/planting). For this visit, growth measurements of the resprouts, SLA measurements, and harvest of the aboveground part of the plants (for a second time) to determine the total biomass of the resprouts are described. https://drive.google.com/file/d/1JCcH8FQEyVol9BSDOPHZ9JnVu1DiXlSo/view?usp=sharing
Thesis
Bachelor thesis
Daniel Cabello Blachowicz (2025) ¿Sembrar o plantar? Comparación de métodos de revegetación en dos quercíneas con diferente ecología. Bachelor thesis, Universidad de Alcalá. Spain.
Supervisor: Pedro Villar-Salvador.
Magazines
Norwegian forestry magazine "Skog"
Hanssen, K.H. and Fløistad, I.S., 2025. Plante eller så eik? Hva gir best resultat? (Planting or sowing of oak - what works best?). In Norwegian. Magasinet Skog no. 2, pp. 20-22.
You can read it here: https://drive.google.com/file/d/16G0vdbzYAyKLuDSCeIm-SDxILqsVHT-E/view?usp=sharing